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Hadrianus (genus) : ウィキペディア英語版 | Hadrianus (genus)
''Hadrianus'' is an extinct genus of tortoise belonging to the Testudinidae〔 found in the United States and Spain and believed to be the oldest true tortoise known.〔 2004 "Skeletochronology as a method of aging Oligocene ''Gopherus laticuneus'' and ''Stylemys nebrascensis'', using ''Gopherus polyphemus'' as a modern analog" Thesis, University of Florida.〕 The genus is thought to be closely related to the genus ''Manouria''.〔 The genus may have evolved in the subtropics of Asia and subsequently migrated to North America and Europe.〔 ==References==
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